LeGros was born in Minneapolis to a teacher mother and real estate broker father and was raised in Redlands, California. He attended the Professional Conservatory at South Coast Repertory in Costa Mesa, California as well as the University of California, Irvine. LeGros is the son-in-law of actor Robert Loggia.
One of his best known roles was in Living in Oblivion in 1995. The film was directed by Tom DiCillo and also starred Steve Buscemi and Catherine Keener. LeGros played Chad Palomino, a male acting diva with endless 'a-list' star demands for a 'b-movie' director and crew.
In 1998 he guest starred in ER as Dr. Max Rocher for three episodes.
From 2000 to 2001 he was a starring cast member on the television show Ally McBeal.
LeGros starred in the Showtime drama Sleeper Cell as Special Agent Ray Fuller in 2005. He also guest starred in the Law & Order episode "Red Ball" in 2005.
He has guest starred in Roseanne, Punky Brewster, The Outer Limits (new series) and Friends.
He portrayed Dr. Dan Harris on the NBC series Mercy.
LeGros portrayed Peter Gray in the Dark Sky thriller Bitter Feast.
In 2011 he starred in the HBO mini-series Mildred Pierce. Also in 2011 he began a recurring role in the FX crime drama Justified. He guest starred as Harlan County criminal Wade Messer in the second season episode "Bloody Harlan". He reprised the role in the third season, appearing in the episode "Harlan Roulette". He returned two years later and reprised his role for the first four episodes of the fifth season. Surprisingly enough, LeGros was also the first actor to portray Raylan Givens in the 1997 TV adaption of the Elmore Leonard novel, Pronto.
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